Nonequilibrium Ecology

The idea that nature is in balance and is only occasionally disrupted by external influences is a central concept in biology. However, much evidence contradicts this assumption. Using evidence from fossil history, studies of plant and animal populations and communities, this book shows that nonequilibrium is more widespread than equilibrium.

Ecology has long been shaped by ideas that stress the sharing of resources and the competition for those resources, and by the assumption that populations and communities typically exist under equilibrium conditions in habitats saturated with both individuals and species. However, much evidence contradicts these assumptions and it is likely that nonequilibrium is much more widespread than might be expected. This book is unique in focusing on nonequilibrium aspects of ecology, providing evidence for nonequilibrium and equilibrium in populations (and metapopulations), in extant communities and in ecological systems over evolutionary time, including nonequilibrium due to recent and present mass extinctions. The assumption that competition is of overriding importance is central to equilibrium ecology, and much space is devoted to its discussion. As communities of some taxa appear to be shaped more by competition than others, an attempt is made to find an explanation for these differences.